Gene Details:

Functional Descriptions:

  • We functionally characterized two genes encoding atypical basic-helix–loop–helix (bHLH) transcription factors, bHLH30 and its homolog bHLH155, which are highly expressed in the elongated adaxial cells.
  • Genetic analyses revealed that bHLH30 and bHLH155 positively regulate LA expansion, and molecular experiments demonstrated their ability to activate the transcription of genes involved in cell elongation and lignification of SCs.
  • Identified two atypical basic-helix–loop–helix (bHLH) transcription factors, bHLH30 and its homolog bHLH155, that are specifically expressed in the ligular region and associate with the promoter regions of maize alpha expansin 5 (EXPA5) and phenylalanine ammonia lyase 7 (PAL7), activating their transcription and thereby promoting LA expansion.
  • Maize bHLH30 and bHLH155 positively regulate LA expansion.

Literature:

Gene Resources:

Orthologs:

Sequences:

cDNA Sequence
  • >Zm00001eb193160_T001 CGGCACACTCCGAGATTCCCCTCAAGGCACCCCAAACCGATGCTTTCTGGCCACTGAAAAGCGCACGGGAGAAGAAGCGCCAAGGCAAAGCAAAAATTTCGCGATGAGTCCACCACGACTCCAAGCAGCTGCGGCCGCGCCGCTCTAACCGGAGCTCCGTTAAGACTCCGCCGCCCCAAATCTCAGCGCCTGAGTCCGCCGCTCTCCTCCCGTGCTCGCCGCCACTGGTTTCCGCGCCGGGACTTGGGTTCGGTGGGGCGACGTGGAGCTTAGGGTTTCGGCAGGATGAGCTACCGGAAGGGGTTGAAGGTTTGGGTGGAGGAGAAGGGGGAAGGGTGGGTGGAGGCGGAGGTGGTGGAAGCGAAGGAGCGCGCCGTCGTCGTGTTCTCGAGCCAGCGGAAAAAGATTACAGTTTCACCAGAAAAATTGTTGCCCAGGGATACAGATGAAGATCTGGGTGGGGGTCATGTGGATGACATGACCAAATTGACATATCTAAATGAACCAGGTGTTCTTTACAATCTGAAGAAAAGATATGCATTGAATGAGATATATACGTACACCGGAAGCATCTTGATTGCCGTTAATCCTTTTACAAGGCTGCCTCACCTGTACAATGAATACATGATGGAACAATACAAGGGCATCCGATTAGGGGAGCTGAGTCCGCATGTTTTTGCGGTGGCGGATGCATCATACAGGGCTATGGTGAATGATTCTCGAAGCCAATCCATCCTGGTTAGTGGTGAAAGTGGTGCTGGCAAAACCGAAACAACGAAGCTTATTATGCAGTATCTCACCTTTGTTGGCGGTAGAGCCGCTCTTGATGACCGGACGGTTGAACAACAGGTTCTAGAATCTAACCCTCTTTTGGAGGCATTTGGCAATGCAAAAACAGTACGCAATGACAATTCAAGTCGATTTGGAAAATTCGTAGAGATACAGTTTGATTCAAGTGGTAGAATATCTGGAGCTGCTATTAGGACCTATCTTTTGGAAAGGTCCCGTGTTGTGCAAATTACTGATCCTGAAAGGAATTTTCATTGCTTCTACCAATTGTGTGCCTCTGGAAAGGACGCAGAGCTGTACAAACTTGGGCATATAAGCAGTTTTCATTACTTGAATCAAAGCAATACACATGATTTGGAAGGGACAAACAATGAAGATGAGTACTGGAAGACAAAAAGGGCAATGGATATTGTTGGGATTAGCAGAGAAGATCAGGACGCTATATTTCGGACATTGGCAGCCATTCTTCATCTTGGCAATATTGAGTTTGTCCCAGGAAAAGATGCTGATTCTTCAAAAATTAAGGATTCAACCTCAAATTTTCATCTCCAAACAGCAGCTAAATTATTTATGTGTGATTCAGATCTATTGGTCTCAACGCTATGTAGCCGATCTATACATACTCGTGAAGGAATAATTGTTAAAGCATTGGATTGTGCTGCTGCCGCAGCTAATCGTGATGCTTTAGCGAAGACTGTATATGCAAGATTATTTGACTGGCTTGTTGAGAATATTAATAAGTCTATTGGTCAAGATGTGGATTCCAAGCTACAAATTGGTGTGCTGGATATCTACGGCTTTGAAAGCTTCAAAAACAACAGCTTTGAGCAGTTCTGCATTAACTTTGCAAATGAAAAACTTCAGCAACACTTTAATGAGCATGTATTCAAGATGGAACAGGAGGAGTACAAAAGTGAGGAAATTAATTGGAGCTACATTGAGTTCATTGACAATCAGGATGTGCTGGATTTGATTGAAAAGAAACCAATTGGAATTATTGCACTGCTGGATGAAGCCTGTATGTTTCCCAAATCTACGCATGAGACCTTTGCGACAAAGATGTTCAGAAACTTCTCTTCCCATCTTAGGCTCGAGAGGACAAAATTCTCAGAAACAGATTTTACCATCTCACACTATGCTGGGAAGGTCACGTATCAAACAGATTCTTTTCTGGAAAAGAATCGAGATTATATTGTTGCAGAACACTGCAATCTTCTATCATCTTCAAGGTGCCCATTTGTTTCTGGACTTTTCACTTCATTGCCTGAAGAATCTATAAGATCTTCATACAAGTTCTCATCAGTTGCTTCCAGATTCAAGCTACAATTGCAAGCCCTGATGGAAACCCTAAACTCGACTGAACCTCACTATGTACGCTGTGTGAAGCCCAACTCTGCTAATCGACCTCAGCTTTTTGAAAATCAAAGTGTATTGCATCAACTTCGTTGTGGGGGTGTCTTAGAGGCTGTCCGTATTAGTCTTGCTGGCTATCCTACCAGAAGGACTTATGCTGAATTTGTTGACCGGTTTGCTGTTCTTGTGCCAGAATTGATGATTGGAAGTTACGATGAAAAAATGATGACAAAGGGTATTCTCGAAAAGATGAAGCTTGAAAATTTTCAACTTGGTAAAACAAAGGTGTTCCTTAGGGCTGGTCAAATTGCGATTTTAGATATGCGCCGTGCTGAGATTTTGGACAACGCTGCACGGCACATTCAAGGCCGTTTCAGAACATTTATCACACGCAAAGAATTTGTGAAAACAAGGGAGGCTTCAATTTCTATACAAGCATACTGCAGAGGCTGTTTGGCAAGGAAGATGTTCGCAAACAGACGAGAAACAGCAGCAGCTGTAATTGTTCAGAAGTATGTCCGAAGATGGCTGCTACGTAGAGCTCACCTGCAAGCATGTTTGGCAGCTCTCCTTATTCAGTCTTACATTCGAGGGTTCATCGCTCGCCGCTATTTCTCTGTCATTAGGGAACATAAAGCTGCTACAGTGATACAGTCCACATGGAGAAGACGGAAGTTTGTCATACTTTTCCAGAATTACCGGCAAGCCACTGTGGCAATTCAGTGTTCTTGGAGACAGAAGCTTGCAAGAAAGGAGCTAAGGAAACTCAAAATGGCTGCAAATGAAGCTGGTGCACTGCGTGAGGCAAAGAATAAGCTTGAGAAAAAGATGGATGATCTTGCTTTAAGACTTACCCTTGAAAGGAGACTGCGGGCTTCCAGTGAAGAGTCAAAATCAGTAGAAATATTGAAGCGTGACAAAATAATTGAATCATTAAGTGCAGAATGTGCTGCCGCTAAGTCAGCTGCTCAAAATGAACATGCCAAAAAATTGCTGCTCCAAAAGCAGTTGGATGATTCTTTGAGAGAGATTACTATGTTGCAGAGTAAAAAGATTATGTCAGCAGAAGCAGCAGAAGAAAACTCCAACCTAAAGAACTTAGTTGAATCATTATCTACGAAGAACTCAATACTGGAGAATGAACTTATTGTGACTCGCAAAAGTAGTGATGATACAATGGAAAAATTGAAGGAAGTCGAGGGGAAATGCAACCATCTACAGCAAAACTTGGACAAATTGCAGGAGAAACTTACAAACTTGGAAAATGAAAATCATGTCTTAAGACAGAAGGCATTTAATATGCCGACAATGAACAATCTGTCTGTAGCTCCAAAGACATTATCTGAGAAATTTTCTGCATCAATTGGGCTTCCCAACAGTGAGCCAAAGCATATATATGAAAGCCCAACACCTACAAAATATCTGGCTTCTCTTCCACAGACTTTGAGTACATCAAGGAGATCGAGGTTGCCTGTTGAAAGGCATGAGCAAAATCATGAAATTTTATTGAGATGCATCAAAGAAAATTTAGGATATAAGGATGGCAAACCAGTTGCAGCATGTATCATTTACAAATGTCTTTTGCACTGGCGTGCTTTTGAATCAGAGCGAACTGCTATTTTTGATCACGTGATTGAGGCCATAAATGATGTCCTAAAGGGGAACGAAGCTGATGGTAGATTACCTTATTGGTTGTCCAATACCTCTGCATTGCTATGCCTTCTCCAAAGAAATCTGCGGTCAAATGGATTATTCACTACCCCGTCTCGTCGTTCTGGTGGAGCTCTAGGGAAGATAGCACAAACATTGAGGTCTCCTTCAAAGTTTATAGGCCGCAGTGATACTCTACCACATGTTGATGCACGGTATCCAGCCATACTATTCAAACAGCAGCTTACTGCATGTGTCGAAAAGATATTTGGGCAGCTCAGAGATAATCTAAAAAAGGAAATATCACCTCTTCTTAATGTTTGCATTCAGGCTCCAAAGTCAACACGTGGGCAATCTGGAAAAGCATCCAAGTCATCTGGAGTTGGTGCCCACCCAGCATCGAACTCCAACTGGGACAATATCGTCAATTTCCTTGACTTACTTATGGATACATTGCGCGAAAATTATGTGCCATCCTTCTTTATACGTAAACTTATCACACAATTATTCTCCTTCATCAATATACAGTTGTTTAACAGTCTTCTTCTCCGACGTGAATGTTGTACATTCTCCAATGGGGAATACGTGAAAGCTGGGCTATCGTTGTTGGAGAAATGGATTACTGATGTCACAGATGAGTTTGCGGGTACATCTTGGCACGAGCTCAATTATATCAGACAAGCTGTTGGATTTTTGGTTATACACCAAAAAAGAAAAAAGACGCTTGAGGAGATCAAGCAAGATCTTTGCCCGTCCTTGAGTGTTCGCCAAATCTATAGAATATGCTCGATGTATTGGGATGACAAATACGGTACCCAAGGCATTTCAACCGAGGTGGTTGCTGCAATGCGGGAGATGGTCAACAAAGATACCCAGAATCTTGTGTCAAATTCCTTCTTGTTGGATGATGACTTAAGTATACCATTCTCTACCGAGGATCTATCAATGGCAATCCCGTCAATAGATTATGCTGACGTCGATCTTCCAGAATCTCTTCAGCACTATACATCAGTACAGTTTCTACTAAGGCAGCAGGATCCACAGCCTGCCCAATAGAATGAATGTGGTCAGCCTGCCCTGAAGATCTGTCTGTCCCCTGGTTGTTTCTACCAAGGCAGCAGGATCCACAGCCTGCGGCCTGCCCAATAGAATGGATGTGGTCAGCCTGCCCTGAAGATCTGTCTGTCCCCTGGTTTTATTGGGGTAACATCCATCTGCGATGTTGTATTGATGTTTAGAGAGTACCTAGTTTGGTGGAGTGGACACAATTTAAAAGCAACCATTGTTGCTAGATGCCAATTCATTGGTGTTCACTCCCCAAGTATTTTTCATTGTACAGGAAATGTTGTAAGTCTGTTGACTAGCGCAAAATTAGTTATATCAGTGGTAAATTGGGCAATGCGCATCAGTGCATC
CDS Sequence
  • >Zm00001eb193160_T001 ATGAGCTACCGGAAGGGGTTGAAGGTTTGGGTGGAGGAGAAGGGGGAAGGGTGGGTGGAGGCGGAGGTGGTGGAAGCGAAGGAGCGCGCCGTCGTCGTGTTCTCGAGCCAGCGGAAAAAGATTACAGTTTCACCAGAAAAATTGTTGCCCAGGGATACAGATGAAGATCTGGGTGGGGGTCATGTGGATGACATGACCAAATTGACATATCTAAATGAACCAGGTGTTCTTTACAATCTGAAGAAAAGATATGCATTGAATGAGATATATACGTACACCGGAAGCATCTTGATTGCCGTTAATCCTTTTACAAGGCTGCCTCACCTGTACAATGAATACATGATGGAACAATACAAGGGCATCCGATTAGGGGAGCTGAGTCCGCATGTTTTTGCGGTGGCGGATGCATCATACAGGGCTATGGTGAATGATTCTCGAAGCCAATCCATCCTGGTTAGTGGTGAAAGTGGTGCTGGCAAAACCGAAACAACGAAGCTTATTATGCAGTATCTCACCTTTGTTGGCGGTAGAGCCGCTCTTGATGACCGGACGGTTGAACAACAGGTTCTAGAATCTAACCCTCTTTTGGAGGCATTTGGCAATGCAAAAACAGTACGCAATGACAATTCAAGTCGATTTGGAAAATTCGTAGAGATACAGTTTGATTCAAGTGGTAGAATATCTGGAGCTGCTATTAGGACCTATCTTTTGGAAAGGTCCCGTGTTGTGCAAATTACTGATCCTGAAAGGAATTTTCATTGCTTCTACCAATTGTGTGCCTCTGGAAAGGACGCAGAGCTGTACAAACTTGGGCATATAAGCAGTTTTCATTACTTGAATCAAAGCAATACACATGATTTGGAAGGGACAAACAATGAAGATGAGTACTGGAAGACAAAAAGGGCAATGGATATTGTTGGGATTAGCAGAGAAGATCAGGACGCTATATTTCGGACATTGGCAGCCATTCTTCATCTTGGCAATATTGAGTTTGTCCCAGGAAAAGATGCTGATTCTTCAAAAATTAAGGATTCAACCTCAAATTTTCATCTCCAAACAGCAGCTAAATTATTTATGTGTGATTCAGATCTATTGGTCTCAACGCTATGTAGCCGATCTATACATACTCGTGAAGGAATAATTGTTAAAGCATTGGATTGTGCTGCTGCCGCAGCTAATCGTGATGCTTTAGCGAAGACTGTATATGCAAGATTATTTGACTGGCTTGTTGAGAATATTAATAAGTCTATTGGTCAAGATGTGGATTCCAAGCTACAAATTGGTGTGCTGGATATCTACGGCTTTGAAAGCTTCAAAAACAACAGCTTTGAGCAGTTCTGCATTAACTTTGCAAATGAAAAACTTCAGCAACACTTTAATGAGCATGTATTCAAGATGGAACAGGAGGAGTACAAAAGTGAGGAAATTAATTGGAGCTACATTGAGTTCATTGACAATCAGGATGTGCTGGATTTGATTGAAAAGAAACCAATTGGAATTATTGCACTGCTGGATGAAGCCTGTATGTTTCCCAAATCTACGCATGAGACCTTTGCGACAAAGATGTTCAGAAACTTCTCTTCCCATCTTAGGCTCGAGAGGACAAAATTCTCAGAAACAGATTTTACCATCTCACACTATGCTGGGAAGGTCACGTATCAAACAGATTCTTTTCTGGAAAAGAATCGAGATTATATTGTTGCAGAACACTGCAATCTTCTATCATCTTCAAGGTGCCCATTTGTTTCTGGACTTTTCACTTCATTGCCTGAAGAATCTATAAGATCTTCATACAAGTTCTCATCAGTTGCTTCCAGATTCAAGCTACAATTGCAAGCCCTGATGGAAACCCTAAACTCGACTGAACCTCACTATGTACGCTGTGTGAAGCCCAACTCTGCTAATCGACCTCAGCTTTTTGAAAATCAAAGTGTATTGCATCAACTTCGTTGTGGGGGTGTCTTAGAGGCTGTCCGTATTAGTCTTGCTGGCTATCCTACCAGAAGGACTTATGCTGAATTTGTTGACCGGTTTGCTGTTCTTGTGCCAGAATTGATGATTGGAAGTTACGATGAAAAAATGATGACAAAGGGTATTCTCGAAAAGATGAAGCTTGAAAATTTTCAACTTGGTAAAACAAAGGTGTTCCTTAGGGCTGGTCAAATTGCGATTTTAGATATGCGCCGTGCTGAGATTTTGGACAACGCTGCACGGCACATTCAAGGCCGTTTCAGAACATTTATCACACGCAAAGAATTTGTGAAAACAAGGGAGGCTTCAATTTCTATACAAGCATACTGCAGAGGCTGTTTGGCAAGGAAGATGTTCGCAAACAGACGAGAAACAGCAGCAGCTGTAATTGTTCAGAAGTATGTCCGAAGATGGCTGCTACGTAGAGCTCACCTGCAAGCATGTTTGGCAGCTCTCCTTATTCAGTCTTACATTCGAGGGTTCATCGCTCGCCGCTATTTCTCTGTCATTAGGGAACATAAAGCTGCTACAGTGATACAGTCCACATGGAGAAGACGGAAGTTTGTCATACTTTTCCAGAATTACCGGCAAGCCACTGTGGCAATTCAGTGTTCTTGGAGACAGAAGCTTGCAAGAAAGGAGCTAAGGAAACTCAAAATGGCTGCAAATGAAGCTGGTGCACTGCGTGAGGCAAAGAATAAGCTTGAGAAAAAGATGGATGATCTTGCTTTAAGACTTACCCTTGAAAGGAGACTGCGGGCTTCCAGTGAAGAGTCAAAATCAGTAGAAATATTGAAGCGTGACAAAATAATTGAATCATTAAGTGCAGAATGTGCTGCCGCTAAGTCAGCTGCTCAAAATGAACATGCCAAAAAATTGCTGCTCCAAAAGCAGTTGGATGATTCTTTGAGAGAGATTACTATGTTGCAGAGTAAAAAGATTATGTCAGCAGAAGCAGCAGAAGAAAACTCCAACCTAAAGAACTTAGTTGAATCATTATCTACGAAGAACTCAATACTGGAGAATGAACTTATTGTGACTCGCAAAAGTAGTGATGATACAATGGAAAAATTGAAGGAAGTCGAGGGGAAATGCAACCATCTACAGCAAAACTTGGACAAATTGCAGGAGAAACTTACAAACTTGGAAAATGAAAATCATGTCTTAAGACAGAAGGCATTTAATATGCCGACAATGAACAATCTGTCTGTAGCTCCAAAGACATTATCTGAGAAATTTTCTGCATCAATTGGGCTTCCCAACAGTGAGCCAAAGCATATATATGAAAGCCCAACACCTACAAAATATCTGGCTTCTCTTCCACAGACTTTGAGTACATCAAGGAGATCGAGGTTGCCTGTTGAAAGGCATGAGCAAAATCATGAAATTTTATTGAGATGCATCAAAGAAAATTTAGGATATAAGGATGGCAAACCAGTTGCAGCATGTATCATTTACAAATGTCTTTTGCACTGGCGTGCTTTTGAATCAGAGCGAACTGCTATTTTTGATCACGTGATTGAGGCCATAAATGATGTCCTAAAGGGGAACGAAGCTGATGGTAGATTACCTTATTGGTTGTCCAATACCTCTGCATTGCTATGCCTTCTCCAAAGAAATCTGCGGTCAAATGGATTATTCACTACCCCGTCTCGTCGTTCTGGTGGAGCTCTAGGGAAGATAGCACAAACATTGAGGTCTCCTTCAAAGTTTATAGGCCGCAGTGATACTCTACCACATGTTGATGCACGGTATCCAGCCATACTATTCAAACAGCAGCTTACTGCATGTGTCGAAAAGATATTTGGGCAGCTCAGAGATAATCTAAAAAAGGAAATATCACCTCTTCTTAATGTTTGCATTCAGGCTCCAAAGTCAACACGTGGGCAATCTGGAAAAGCATCCAAGTCATCTGGAGTTGGTGCCCACCCAGCATCGAACTCCAACTGGGACAATATCGTCAATTTCCTTGACTTACTTATGGATACATTGCGCGAAAATTATGTGCCATCCTTCTTTATACGTAAACTTATCACACAATTATTCTCCTTCATCAATATACAGTTGTTTAACAGTCTTCTTCTCCGACGTGAATGTTGTACATTCTCCAATGGGGAATACGTGAAAGCTGGGCTATCGTTGTTGGAGAAATGGATTACTGATGTCACAGATGAGTTTGCGGGTACATCTTGGCACGAGCTCAATTATATCAGACAAGCTGTTGGATTTTTGGTTATACACCAAAAAAGAAAAAAGACGCTTGAGGAGATCAAGCAAGATCTTTGCCCGTCCTTGAGTGTTCGCCAAATCTATAGAATATGCTCGATGTATTGGGATGACAAATACGGTACCCAAGGCATTTCAACCGAGGTGGTTGCTGCAATGCGGGAGATGGTCAACAAAGATACCCAGAATCTTGTGTCAAATTCCTTCTTGTTGGATGATGACTTAAGTATACCATTCTCTACCGAGGATCTATCAATGGCAATCCCGTCAATAGATTATGCTGACGTCGATCTTCCAGAATCTCTTCAGCACTATACATCAGTACAGTTTCTACTAAGGCAGCAGGATCCACAGCCTGCCCAATAG
Protein Sequence
  • >Zm00001eb193160_P001 MSYRKGLKVWVEEKGEGWVEAEVVEAKERAVVVFSSQRKKITVSPEKLLPRDTDEDLGGGHVDDMTKLTYLNEPGVLYNLKKRYALNEIYTYTGSILIAVNPFTRLPHLYNEYMMEQYKGIRLGELSPHVFAVADASYRAMVNDSRSQSILVSGESGAGKTETTKLIMQYLTFVGGRAALDDRTVEQQVLESNPLLEAFGNAKTVRNDNSSRFGKFVEIQFDSSGRISGAAIRTYLLERSRVVQITDPERNFHCFYQLCASGKDAELYKLGHISSFHYLNQSNTHDLEGTNNEDEYWKTKRAMDIVGISREDQDAIFRTLAAILHLGNIEFVPGKDADSSKIKDSTSNFHLQTAAKLFMCDSDLLVSTLCSRSIHTREGIIVKALDCAAAAANRDALAKTVYARLFDWLVENINKSIGQDVDSKLQIGVLDIYGFESFKNNSFEQFCINFANEKLQQHFNEHVFKMEQEEYKSEEINWSYIEFIDNQDVLDLIEKKPIGIIALLDEACMFPKSTHETFATKMFRNFSSHLRLERTKFSETDFTISHYAGKVTYQTDSFLEKNRDYIVAEHCNLLSSSRCPFVSGLFTSLPEESIRSSYKFSSVASRFKLQLQALMETLNSTEPHYVRCVKPNSANRPQLFENQSVLHQLRCGGVLEAVRISLAGYPTRRTYAEFVDRFAVLVPELMIGSYDEKMMTKGILEKMKLENFQLGKTKVFLRAGQIAILDMRRAEILDNAARHIQGRFRTFITRKEFVKTREASISIQAYCRGCLARKMFANRRETAAAVIVQKYVRRWLLRRAHLQACLAALLIQSYIRGFIARRYFSVIREHKAATVIQSTWRRRKFVILFQNYRQATVAIQCSWRQKLARKELRKLKMAANEAGALREAKNKLEKKMDDLALRLTLERRLRASSEESKSVEILKRDKIIESLSAECAAAKSAAQNEHAKKLLLQKQLDDSLREITMLQSKKIMSAEAAEENSNLKNLVESLSTKNSILENELIVTRKSSDDTMEKLKEVEGKCNHLQQNLDKLQEKLTNLENENHVLRQKAFNMPTMNNLSVAPKTLSEKFSASIGLPNSEPKHIYESPTPTKYLASLPQTLSTSRRSRLPVERHEQNHEILLRCIKENLGYKDGKPVAACIIYKCLLHWRAFESERTAIFDHVIEAINDVLKGNEADGRLPYWLSNTSALLCLLQRNLRSNGLFTTPSRRSGGALGKIAQTLRSPSKFIGRSDTLPHVDARYPAILFKQQLTACVEKIFGQLRDNLKKEISPLLNVCIQAPKSTRGQSGKASKSSGVGAHPASNSNWDNIVNFLDLLMDTLRENYVPSFFIRKLITQLFSFINIQLFNSLLLRRECCTFSNGEYVKAGLSLLEKWITDVTDEFAGTSWHELNYIRQAVGFLVIHQKRKKTLEEIKQDLCPSLSVRQIYRICSMYWDDKYGTQGISTEVVAAMREMVNKDTQNLVSNSFLLDDDLSIPFSTEDLSMAIPSIDYADVDLPESLQHYTSVQFLLRQQDPQPAQ